Lemurs of Madagascar
Madagascar is world-famous for its lemurs—primates that look something like a cat crossed with a squirrel and a dog. These animals are unique to the island and display a range of interesting behaviors from singing like a whale (the indri) to sashaying across the sand like a ballet dancer (the sifaka). Below you will learn more about these fascinating creatures.
Lemur history
Madagascar lacks the dominant form of primate distributed worldwide, those of the suborder Haplorhini (monkeys, chimps, gorillas, and Homo sapiens). Instead, their niche has been filled by an older group of primates, the lemurs. Lemurs belong to the sub-order Strepsirhini together with bushbabies, lorises, and pottos which—like the original lemurs—are nocturnal, insectivorous primates characterized by a small body, a long nose, and large eyes. Lemurs have an interesting evolutionary history and the only reason they still exist today is because of Madagascar's isolation.
Until around 160 million years ago, Madagascar was attached to the African mainland as part of the super continent Gondwanaland (formed of Africa, South America, Australia, Antarctica, India, and Madagascar). As Gondwanaland broke apart, Madgascar moved away from Africa. The first lemur-like primates on the fossil record appeared roughly 60 million years ago in mainland Africa and crossed over to Madagascar shortly thereafter.
The island continued to drift eastward and by the time monkeys appeared on the scene 17-23 million years ago, Madagascar was isolated from their arrival. As highly intelligent and adaptive primates, monkeys quickly drove the lemur lineage elsewhere in the world toward extinction (a few Strepsirhines—including bushbabies, lorises, and pottos—managed to hang on by retaining their nocturnal, solitary, and insectivorous traits).
Madagascar's lemurs—isolated from evolutionary changes of the world—radiated into the large island's many niches without much competition or predation. Today lemurs are found in virtually all of Madagascar's ecosystems and share some of the social and behavioral characteristics of monkeys (i.e., forming social groups, eating fruit and vegetation, and being active during the day).
Upper primates did not reach Madagascar until about they learned to navigate the high seas and arrived on boats roughly 2,000 years ago. Humans quickly went to work on the island's lemurs, reducing the number of species found in Madagascar by at least 15. The largest species suffered the most and today the largest remaining lemur is the Indri which would have been dwarfed by the gorilla-sized species once found on the island. Currently nearly all lemurs are endangered species, due mainly to habitat destruction (deforestation) and hunting.
Lemurs today
Today Madagascar is home to over 110 species of lemurs across five families and 14 genera ranging in size from the 25-gram pygmy mouse lemur to the indri. All these species are endemic to Madagascar (two lemur species were introduced to the Comoros) giving the country the highest number of primate species (Brazil, which has 77 species but only two endemic genera and no endemic families, is second). And new species are still being discovered— between 2000 and 2008, 39 new species were described.
Global importance of Madagascar's lemurs
According to Russell Mittermeier in The Eighth Continent, although Madagascar "is only one of 92 countries with wild primate populations, it is alone responsible for 21 percent (14 of 65) of all primate genera and 36 percent (five of 14) of all primate families, making it the single highest priority" for primate conservation. "Madagascar is so important for primates that primatologists divide the world into four major regions: the whole of South and Central America, all of southern and southeast Asia, mainland Africa, and Madagascar, which ranks as a full-fledged region all by itself."
Behavior
Non-scientists generally group lemurs by their primary time of activity: day or night. Nocturnal lemurs are typically smaller and more reclusive than their diurnal counterparts. Lemurs are vocal animals, making sounds that range from the grunts and swears of brown lemurs and sifaka to the chirps of mouse lemurs to the eerie, wailing call of the indri, which has been likened to a cross between a police siren and the song of a humpback whale.
Lemur photos
Lemur News
The fuel that moves people: the Ecuadorian case
- In Ecuador, the main areas of colonization were a north-south corridor along the base of the Andes and the Sucumbíos-Orellana quadrant, the country’s major oil-producing region.
- Since the 1970s, populations in both areas have grown significantly. The Andean zone went from 160,000 inhabitants to more than 520,000 in 2017; in parallel, the population in the provinces of Sucumbíos and Orellana increased from less than 12,000 to more than 350,000.
- Colonization also led to the invasion of lands of the indigenous Shuar, which prompted an unusual effort on their part to protect their territory. Today, the area specializes in cattle production and seeks to establish a niche market for high-quality beef for the domestic market.
Brazil’s illegal gold miners carve out new Amazon hotspots in conservation units
- President Luiz Inácio Lula da Silva’s administration reduced the expansion of illegal gold mining in the Brazilian Amazon, but miners keep finding new sites.
- In 15 conservation units, illegal gold miners destroyed 330 hectares (815 acres) in only two months.
- According to experts, gold miners expelled from Indigenous territories may be migrating to conservation units.
- Alliances with narco mafias and the rise in gold prices are obstacles to fighting illegal mining.
Amazon’s Boiling River gives scientists a window into the rainforest’s future
- Scientists studying Peru’s Boiling River found 11% fewer tree species for every 1°C (1.8°F) increase in temperature, offering insights into how climate change might affect the Amazon Rainforest.
- The research team discovered that hotter areas not only had fewer species overall but were dominated by heat-tolerant trees that typically grow in the warmest parts of South America.
- The study site is protected by Indigenous Asháninka people as sacred land, but the forest still faces threats from nearby deforestation and fires, reflecting broader challenges across the Amazon.
- The Amazon is experiencing climate pressures, with fire-affected areas in the Brazilian Amazon increasing 18-fold in September 2024, covering a combined area nearly the size of the Netherlands.
Electrochemical removal of ocean CO2 offers potential — and concerns
- Stripping seawater of carbon dioxide via electrochemical processes — thereby prompting oceans to draw down more greenhouse gas from the atmosphere — is a geoengineering approach under consideration for largescale CO2 removal. Several startups and existing companies are planning projects at various scales.
- Once removed from seawater, captured carbon dioxide can be stored geologically or used commercially by industry. Another electrochemical method returns alkaline seawater to the oceans, causing increased carbon dioxide absorption over time.
- In theory, these techniques could aid in carbon emission storage. But experts warn that as some companies rush to commercialize the tech and sell carbon credits, significant knowledge gaps remain, with potential ecological harm needing to be determined.
- Achieving the scale required to make a dent in climate change would require deploying huge numbers of electrochemical plants globally — a costly and environmentally risky scenario deemed unfeasible by some. One problem: the harm posed by scale-up isn’t easy to assess with modeling and small-scale projects.
South Korea slashes forest biomass energy subsidies in major policy reform
- In a surprise move, South Korea has announced that it will end subsidies for all new biomass projects and for existing state-owned plants cofiring biomass with coal, effective January 2025, a significant and sudden policy shift.
- Additionally, government financial support for dedicated biomass plants using imported biomass will be phased down, while support for privately owned cofiring plants will be phased out over the next decade. However, subsidy levels for domestically produced biomass fuel remain unchanged.
- The biomass reform is being hailed by forest advocates as a step in the right direction, potentially setting a new, environmentally sound precedent for the region.
- Advocates are now calling on Japan, Asia’s largest forest biomass importer, to follow South Korea’s example.
Coral destruction for toilet construction: Interview with a Malagasy fisher
- Toamasina, a coastal city in eastern Madagascar, is surrounded by an extensive network of coral reefs that are home to near-threatened species.
- For decades, these reefs have been under threat from an unusual activity: The use of coral in the construction of septic tanks.
- Mongabay spoke with Abraham Botovao, a boat skipper and the president of a local fishers’ association, who has been closely monitoring this trade and its impact on the local marine environment.
- “It frustrates me every time I see them when I’m out fishing, but unfortunately all I can do is watch without being able to do anything,” Botovao said.
Poachers target South Africa’s ‘miracle’ plant with near impunity
- South Africa has faced a surge in poaching of rare succulents by criminal syndicates since 2019.
- A recent spike in prices paid for a different kind of plant, a drylands-adapted lily, the miracle clivia (Clivia mirabilis), has drawn the attention of plant-trafficking syndicates to the lone reserve where it grows.
- Large numbers of clivias have been seized by law enforcement, raising fears that this rare plant is quickly being wiped out from the limited range where it’s known to occur.
- Reserve staff and law enforcement agencies are underfunded and spread too thinly across the vast landscapes of South Africa’s Northern Cape province targeted by plant poachers.
As lithium mining bleeds Atacama salt flat dry, Indigenous communities hit back
- The Council of Atacameño Peoples filed a complaint in October 2024 against lithium mining companies operating in Chile’s Atacama salt flat, accusing them of causing the land to sink around their extraction wells.
- The complaint was based on findings from a study published in July that revealed portions of the salt flat are subsiding by up to 2 centimeters, or nearly an inch, per year.
- Scientists warn that one of the main consequences could be the loss of the aquifer’s storage capacity.
- They also point out that since the salt flat lies on a tectonic fault, the subsidence could spread further, including to two protected areas in the region that are home to flamingos and other rare wildlife.
Indonesia’s Indigenous communities sidelined from conservation
- Research shows that globally, Indigenous peoples are the most effective stewards of their forests and the massive stores of carbon and biodiversity within.
- Yet in Indonesia, which harbors the majority of Earth’s species, Indigenous communities are increasingly sidelined from nature conservation efforts.
- Activists say it is urgent for the Indonesian government to pass a long-awaited bill on Indigenous rights to ensure that Indigenous peoples can contribute to biodiversity conservation without fear of being criminalized or evicted.
- This is especially important, activists say, in light of a new conservation law in Indonesia, which is criticized for not protecting Indigenous land rights; the law also outlines a new form of “preservation area,” where Indigenous activities could be heavily restricted.
Balochistan’s Gwadar city sits at the crossroads of climate and conflict
- A new study examines the links between conflict and climate in Pakistan’s Balochistan province, where extreme weather can be a threat multiplier.
- The port city of Gwadar serves as an example, as local residents have long had grievances against the state, which were exacerbated by recent flooding that killed several people and displaced hundreds.
- Experts highlight the absence of data-driven policies, citing a gap in research that has hindered solutions; they call for investment in data and the inclusion of local people in decision-making and infrastructure planning.
From Bhutan to Nigeria & Kenya, women endure climate change differently than men
- Research shows that globally, women and girls suffer greater effects of climate change and environmental disasters than men; at the same time, women environmental journalists often face greater obstacles on the job, and women’s voices are often missing from stories about climate change.
- Three recent Mongabay fellows, all women, report on specific examples from their home countries (Bhutan, Nigeria, Kenya) in which women disproportionately experience the effects of climate change and extreme weather.
- In all three examples, women exhibit a perseverance that ensures their own and their families’ survival — and sometimes aids their own independence and resourcefulness.
For ecological restoration, evidence-based standards deliver better outcomes (commentary)
- The UN Decade on Ecosystem Restoration has triggered a global movement to rally individual action, financial investment, and political backing to prevent, halt and reverse the loss of nature.
- Evidence-based standards can help meet restoration targets and improve general compliance with laws and regulations while delivering social, environmental and economic net gain for people and nature.
- “As we near the halfway point of the UN Decade on Ecosystem Restoration [the] global application of effective restoration through the use of standards provides a path forward that can help slow climate change and recover ecosystem processes and biodiversity for the future of life on Earth,” a new op-ed argues.
- This article is a commentary. The views expressed are those of the authors, not necessarily Mongabay.
Unlike: Brazil Facebook groups give poachers safe space to flex their kills
- A new study shows how openly poachers in Brazil are sharing content of dead wildlife, including threatened and protected species, on Facebook.
- It found 2,000 records of poaching on Brazilian Facebook groups between 2018 and 2020, amounting to 4,658 animals from 157 species from all over the country.
- Data suggest there were trophy hunts, meant only to show off hunting hauls rather than being done for subsistence or a consequence of human-wildlife conflict.
- The study highlights the impunity for environmental crimes and the easy dissemination of content related to illegal practices on social media networks in Brazil.
‘Killed while poaching’: When wildlife enforcement blurs into violence
- In October 2023, Mongabay traveled to Uganda’s Queen Elizabeth National Park as part of a reporting series on protected areas in East Africa.
- While there, we heard allegations that Uganda Wildlife Authority rangers have carried out extrajudicial killings of suspected bushmeat poachers inside the park.
- Two weeks before our visit, a man was shot to death inside the park; his relatives and local officials alleged he was killed by wildlife rangers while attempting to surrender.
- The allegations follow other recent human rights scandals related to aggressive conservation enforcement practices in the nearby Congo Basin.
Most large banks failing to consider Indigenous rights
- A new report by finance watchdog BankTrack evaluated the policies and practices of 50 major banks and found that most are failing to fully implement adequate safeguards in line with U.N. human rights principles.
- The 2024 report included three new criteria centered around the rights of human rights defenders and Indigenous peoples and the right to a healthy environment; the majority of banks did not explicitly acknowledge environmental rights are human rights and all failed in due diligence around Indigenous peoples’ free, prior and informed consent.
- The report found that small progress has been made in the last two years as banks improve policies and processes for managing human rights.
- The authors say stronger human rights due diligence laws could be a game changer in driving corporate respect for human rights.
New frog species show how geology shapes Amazon’s biodiversity
- DNA testing of two new-to-science frog species has shown they share a common ancestor — a species that lived 55 million years ago in the mountains of what is today Brazil’s Amazonas state.
- The multidisciplinary study drew together biologists and geologists to map how geological changes in the mountain range shaped not just its geography but also the diversity of species in the region.
- The two endemic species were collected on two separate peaks — Neblina and Imeri — and their discovery has led to further understanding of the origins and evolution of biodiversity in the Amazon.
- Another expedition to the Tulu-Tuloi Range, located 200 kilometers (120 miles) from Imeri, is scheduled for 2025.
Even for ‘progressive’ Danone, complying with EUDR is a challenge
- The EU deforestation-free products regulation (EUDR) requires companies importing cocoa, cattle, rubber, soy, wood, palm oil and coffee into the EU to demonstrate their products weren’t grown on land deforested after Dec. 31, 2020.
- French dairy giant Danone works with soy, cocoa and palm oil — products subject to the EUDR, which goes into effect at the end of 2025.
- In general, Danone’s sustainability policies have been better than most, outsider observers said. But it still has a lot of work to do to ensure 100% of its supply chain is free of deforestation and land conversion.
A port is destroying corals to expand. Can an NGO rescue enough to matter?
- The ongoing expansion of the port of Toamasina in eastern Madagascar is set to destroy 25 hectares (62 acres) of coral reefs.
- Tany Ifandovana, a Malagasy NGO, removed a small portion of these corals before construction began, and transplanted them to a coral island several kilometers away, as a way to ecologically compensate for the losses, at least in part.
- The NGO faces major challenges, including a lack of resources, little support from the port, and locals destroying corals around the island transplant site.
- “As an environmentalist, it hurt my heart to know that these corals were just going to be filled in,” Tany Ifandovana’s vice president told Mongabay. “Something had to be done.”
Nepal created a forest fund to do everything; five years on it’s done nothing
- Nepal’s Forest Development Fund, established in 2019, was designed to support forest conservation, research and other environmental initiatives, but it has not spent any of the allocated funds in five years.
- The fund is meant to be financed through various sources, including lease fees from developers, compensatory afforestation payments, a percentage of profits from forest land use and revenue from carbon trading.
- Forest user communities, which have successfully increased forest cover in Nepal, continue to face financial difficulties, with illegal logging and wildfires exacerbating the situation, while the FDF remains frozen.
Can the Cali Fund provide a rights-based remedy for biopiracy? (commentary)
- One ongoing element of wealth extraction from the Global South that remains largely unaddressed – biopiracy – requires a human rights-based response, a new op-ed argues.
- Defined as the unauthorized use of genetic resources and traditional knowledge of Indigenous communities and developing nations for profit without their consent, a remedy to biopiracy was recently agreed to at the UN Biodiversity Conference (COP16) in Colombia.
- Can the Cali Fund – which obliges corporations that profit from biodiversity to contribute to its conservation – be a step in the right direction, the authors ask?
- This article is a commentary. The views expressed are those of the authors, not necessarily Mongabay.
Lemur species counts for selected parks
Protected Area | Species | |
Andringitra | 13 | |
Zahamena | 13 | |
Marojejy | 12 | |
Ranomafana | 12 | |
Andasibe Mantadia | 11 | |
Bemaraha | 11 | |
Marotandrano | 11 | |
Ankarana | 10 | |
Masoala | 10 | |
Kirindy Mitea | 8 | |
Montagne d'Ambre | 8 | |
Namoroka | 8 | |
Zombitse-Vohibasia | 8 | |
Isalo | 7 | |
Kasijy | 7 | |
Manombo | 7 | |
Pic d'Ivohibe | 7 | |
Bay de baly | 6 | |
Bemarivo | 6 | |
Bora | 6 | |
Nosy Mangabe | 6 | |
Tampoketsa-Analamaintso | 3 | |
Tsimanampetsotsa | 3 | |
Ambohijanahary | 2 |
Lemur species
Please note: a number of lemur species have been described since this list was published.
Family | Common name | Scientific name | Local name | Active | Pictures |
Cheirogaleidae | Mouse and Dwarf lemurs | Nocturnal | |||
Cheirogaleidae | Hairy-eared Dwarf Mouse-lemur | Allocebus trichotis | Nocturnal | ||
Cheirogaleidae | Southern Fat-tailed Dwarf Lemur | Cheirogaleus adipicaudatus | Matavirambo | Nocturnal | |
Cheirogaleidae | Furry-eared Dwarf Lemur | Cheirogaleus crossleyi | Matavirambo | Nocturnal | |
Cheirogaleidae | Greater Dwarf Lemur | Cheirogaleus major | Matavirambo | Nocturnal | + |
Cheirogaleidae | Western Fat-tailed Dwarf Lemur | Cheirogaleus medius | Matavirambo, Kely Be-ohy, Tsidy, Tsidihy | Nocturnal | |
Cheirogaleidae | Lesser Iron Gray Dwarf Lemur | Cheirogaleus minusculus | Matavirambo | Nocturnal | |
Cheirogaleidae | Greater Iron Gray Dwarf Lemur | Cheirogaleus ravus | Matavirambo | Nocturnal | |
Cheirogaleidae | Sibree's Dwarf Lemur | Cheirogaleus sibreei | Matavirambo | Nocturnal | |
Cheirogaleidae | Gray Mouse-lemur | Microcebus murinus | Tsidy, Koitsiky, Titilivaha, Vakiandri, Pondiky | Nocturnal | + |
Cheirogaleidae | Pygmy Mouse-lemur | Microcebus myoxinus | Tsidy | Nocturnal | |
Cheirogaleidae | Golden Mouse-lemur | Microcebus ravelobensis | Tsidy | Nocturnal | |
Cheirogaleidae | Red Mouse-lemur | Microcebus rufus | Anakatsidina, Tsidy, Tsitsidy, Tistsihy | Nocturnal | + |
Cheirogaleidae | Giant Mouse-lemur or Coquerel's Mouse-lemur | Mirza coquereli | Tsiba, Tilitilivaha, Siba, Setohy, Fitily | Nocturnal | + |
Cheirogaleidae | Amber Mountain Fork-crowned Lemur | Phaner electromontis | Tanta, Tantaraolana | Nocturnal | |
Cheirogaleidae | Masoala Fork-crowned Lemur | Phaner furcifer | Tanta, Tantaraolana | Nocturnal | |
Cheirogaleidae | Western Fork-crowned Lemur | Phaner pallescens | Tanta, Tantaraolana, Vakivoho | Nocturnal | |
Cheirogaleidae | Sambirano Fork-crowned Lemur | Phaner parienti | Tanta, Tantaraolana | Nocturnal | |
Daubentoniidae | Aye-aye | Nocturnal | |||
Daubentoniidae | Aye-aye | Daubentonia madagascariensis | Aye-aye, Ahay, Itay-hay, Aiay | Nocturnal | |
Indridae | Woolly lemurs and allies | Diurnal | |||
Indridae | Eastern Avahi | Avahi laniger | Avahina, Avahy, Ampongy, Fotsifaka | Nocturnal | + |
Indridae | Western Avahi | Avahi occidentalis | Fotsife, Tsarafangitra | Nocturnal | |
Indridae | Indri lemur | Indri indri indri | Babakoto, Amboanala | Diurnal | + |
Indridae | Indri lemur | Indri indri variegatus | Babakoto, Amboanala | Diurnal | + |
Indridae | Coquerel's Sifaka | Propithecus coquereli | Ankomba malandy, Sifaka, Tsibahaka | Diurnal | |
Indridae | Crowned Sifaka | Propithecus deckenii coronatus | Tsibahaka, Sifaka | Diurnal | |
Indridae | Decken's Sifaka | Propithecus deckenii dekenii | Tsibahaka, Sifaka | Diurnal | + |
Indridae | Silky Sifaka | Propithecus diadema candidus | Simpona, Simpony | Diurnal | |
Indridae | Diademed Sifaka | Propithecus diadema diadema | Simpona, Simpony | Diurnal | + |
Indridae | Milne-Edwards's Sifaka | Propithecus edwardsi | Simpona, Simpony | Diurnal | + |
Indridae | Perrier's Sifaka | Propithecus perrieri | Radjako, Ankomba Job | Diurnal | |
Indridae | Tattersall's Sifaka | Propithecus tattersalli | Ankomba malandy, Simpona | Diurnal | |
Indridae | Verreaux's Sifaka | Propithecus verreauxi | Sifaka | Diurnal | + |
Lemuridae | True lemurs | Diurnal | |||
Lemuridae | White-fronted Lemur | Eulemur albifrons | Varika | Diurnal | |
Lemuridae | White-collared Lemur | Eulemur albocollaris | Varika | Diurnal | |
Lemuridae | Red-collared Lemur | Eulemur collaris | Varika | Diurnal | + |
Lemuridae | Crowned Lemur | Eulemur coronatus | Varika | Diurnal | |
Lemuridae | Brown Lemur | Eulemur fulvus | Varikamavo, Komba | Diurnal | + |
Lemuridae | Black Lemur | Eulemur macaco | Ankomba, Komba | Diurnal | + |
Lemuridae | Blue-eyed Black Lemur | Eulemur macaco flavifrons | Ankomba, Komba | Diurnal | + |
Lemuridae | Mongoose Lemur | Eulemur mongoz | Komba | Diurnal | |
Lemuridae | Red-bellied Lemur | Eulemur rubriventer | Varikamena | Diurnal | |
Lemuridae | Red-fronted Lemur | Eulemur rufus | Varika, Varikamavo | Diurnal | + |
Lemuridae | Sanford's Lemur | Eulemur sanfordi | Ankomba, Beharavoaka | Diurnal | |
Lemuridae | Alaotran Gentle Lemur | Hapalemur alaotrensis | Bandro | Diurnal | |
Lemuridae | Golden Gentle Lemur | Hapalemur aureus | Varibolomena, Bokombolomena | Diurnal | |
Lemuridae | Gray Gentle Lemur | Hapalemur griseus | Varibolomadinika | Diurnal | + |
Lemuridae | Sambriano Gentle Lemur | Hapalemur occidentalis | Bekola, Kofi, Ankomba valiha | Diurnal | |
Lemuridae | Ring-tailed Lemur | Lemur catta | Maki, Hira | Diurnal | + |
Lemuridae | Broad-nosed Gentle Lemur | Prolemur simus | Varibolomavo, Vari, Varikandra | Diurnal | |
Lemuridae | Red Ruffed Lemur | Varecia rubra | Varimena | Diurnal | + |
Lemuridae | Black-and-White Ruffed Lemur | Varecia variegata | Varijatsy | Diurnal | + |
Megaladapidae | Sportive lemurs | Nocturnal | |||
Megaladapidae | Back-striped Sportive Lemur | Lepilemur dorsalis | Apongy | Nocturnal | |
Megaladapidae | Milne-Edwards's Sportive Lemur | Lepilemur edwardsi | Boenga, Boengy, Repahaka | Nocturnal | + |
Megaladapidae | White-footed Sportive Lemur | Lepilemur leucopus | Songiky | Nocturnal | + |
Megaladapidae | Small-toothed Sportive Lemur | Lepilemur microdon | Trangalavaka, Kotrika or Kotreka, Fitiliky, Itataka, Varikosy | Nocturnal | |
Megaladapidae | Mitsinjo Sportive Lemur | Lepilemur mitsinjonensis | Kotrika, Varikosy | Nocturnal | |
Megaladapidae | Weasel Lemur | Lepilemur mustelinus | Trangalavaka, Kotrika, Fitiliky, Itataka, Varikosy | Nocturnal | |
Megaladapidae | Red-tailed Sportive Lemur | Lepilemur ruficaudatus | Boenga, Boengy | Nocturnal | |
Megaladapidae | Ankarana Sportive Lemur | Lepilemur septentrionalis ankaranensis | Mahiabeala, Songiky | Nocturnal | |
Megaladapidae | Seal's Sportive Lemur | Lepilemur seali | Songiky | Nocturnal | |
Megaladapidae | Northern Sportive Lemur | Lepilemur septentrionalis septentrionalis | Mahiabeala, Songiky | Nocturnal |